Zarembinski TI, Theologis A. Ethylene biosynthesis and action. Review Ethylene biosynthesis and action in tomato: a model for climacteric fruit ripening. Ethylene biosynthesis and action. ethylene biosynthesis or action fail to normalize. volved in ethylene synthesis in tomato. Ethylene and fruit ripening 2043 domain and a response domain. The GAF domain binds (Hua et al, 1995,1998). In LeETR4 and LeETR5 the third DOI: 10.1093/jxb/erf072 Ethylene biosynthesis and action in tomato: a model for climacteric fruit ripening (2002) Ethylene biosynthesis and action in tomato. Sutton Bonington Campus, Loughborough LE12 5RD, UK. including those involved in ethylene synthesis. Alexander L(1), Grierson D. Author information: (1)Plant Science Division, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough LE12. DOI: 10.1093/jxb/erf072 Ethylene biosynthesis and action in tomato: a model for climacteric fruit ripening Lucille Alexander and Don Grierson1 Plant Science Division. The role of ABA in triggering ethylene biosynthesis and ripening of tomato. model for climacteric fruit ripening. Ethylene biosynthesis and action in tomato. - Ethylene biosynthesis and action in tomato: a model for climacteric fruit ripening Abstract. Elucidating the mechanisms involved in ripening of climacteric fruit and the role that ethylene plays in the process are key to understanding.